És maga a kiindulási cikk:

Wang et al: High-coverage genome of the Tyrolean Iceman reveals unusually high Anatolian farmer ancestry (2023)

https://www.cell.com/cell-genomics/pdfExtended/S2666-979X(23)00174-X

Kb. a lényeg (bőrszín):

"The weighted genetic score of dark pigmentation in the Iceman is estimated to be 0.591, higher than the score of present-day southern European populations taking Sardinians as an example (Table S11), which the Iceman shares closest genetic affinity to (Figure S1) and which represent the highest level of pigmentation among modern-day European

groups,29 although it is lower than the score of ancient LBK farmers and the Luxembourg_Loschbour.DG hunter-gatherer."